The evolution of language implies the parallel evolution of an ability to respond appropriately to signals (language understanding) and an ability to produce the appropriate signals in the appropriate circumstances (language production). When linguistic signals are produced to inform other individuals, individuals that respond appropriately to these signals may increase their reproductive chances but it is less clear what is the reproductive advantage for the languages producers. We present simulations in which populations of neural networks living in an environment evolve a simple language with an informative function. Signals are produced to help other individuals to categorize edible and poisonous mushrooms in order to decide whether to approach or avoid encountered mushrooms. Language production, while not under direct evolutionary pressure, evolves as a by-product of the independently evolving perceptual ability to categorize mushrooms. Keywords: Language evolution, Ge...

This paper describes different types of models for the evolution of communication and language. It uses the distinction between signals, symbols, and words for the analysis of evolutionary models of language. In particular, it show how evolutionary computation techniques, such as Artificial Life, can be used to study the emergence of syntax and symbols from simple communication signals. Initially, a computational model that evolves repertoires of isolated signals is presented. This study has simulated the emergence of signals for naming foods in a population of foragers. This type of model studies communication systems based on simple signal-object associations. Subsequently, models that study the emergence of grounded symbols are discussed in general, including a detailed description of a work on the evolution of simple syntactic rules. This model focuses on the emergence of symbol-symbol relationships in evolved languages. Finally, computational models of syntax acquisition and evolution are discussed. These different types of computational models provide an operational def'mition of the signal/symbol/word distinction. The simulation and analysis of these types of models will help understanding the role of symbols and symbol acquisition in the origin of language.

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this paper is on procedures whereby new (e.g., juvenile) members of a population could learn to communicate with the other members by observing their communicative behavior. Two apparently distinct issues are relevant to the evaluation of such learning procedures. First, the procedure must enable the new members to accurately acquire the communication system of the population, even though their observations may be limited, noisy, or otherwise misleading. Second, the learning procedure used by its new members will affect the population's communication system over time. The use of a particular procedure might result in the population's communication increasing in coordination, ultimately yielding a nearly optimally coordinated system. If a learning procedure were to satisfy both criteria, it could explain how learned communication systems are maintained over time, as well as how they are established in the first place.

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On the evolutionary trajectory that led to human language there must have been a transition from a fairly limited to an essentially unlimited communication system. The structure of modern human languages reveals at least two steps that are required for such a transition: in all languages (i) a small number of phonemes are used to generate a large number of words; and (ii) a large number of words are used to a produce an unlimited number of sentences. The first (and simpler) step is the topic of the current paper. We study the evolution of communication in the presence of errors and show that this limits the number of objects (or concepts) that can be described by a simple communication system. The evolutionary optimum is achieved by using only a small number of signals to describe a few valuable concepts. Adding more signals does not increase the fitness of a language. This represents an error limit for the evolution of communication. We show that this error limit can be overcome by combining signals (phonemes) into words. The transition from an analogue to a digital system was a necessary step toward the evolution of human language.

The lexical matrix is an integral part of the human language system. It provides the link between word form and word meaning. A simple lexical matrix is also at the center of any animal communication system, where it defines the associations between form and meaning of animal signals. We study the evolution and population dynamics of the lexical matrix. We assume that children learn the lexical matrix of their parents. This learning process is subject to mistakes: (i) children may not acquire all lexical items of their parents (incomplete learning); and (ii) children might acquire associations between word forms and word meanings that differ from their parents ’ lexical items (incorrect learning). We derive an analytic framework that deals with incomplete learning. We calculate the maximum error rate that is compatible with a population maintaining a coherent lexical matrix of a given size. We calculate the equilibrium distribution of the number of lexical items known to individuals. Our analytic investigations are supplemented by numerical simulations that describe both incomplete and incorrect learning, and other extensions. c ○ 2001 Society for Mathematical Biology 1.

John Maynard Smith and Eörs Szathmáry argued that human language signified the eighth major transition in evolution: human language marked a new form of information transmission from one generation to another [Maynard Smith J, Szathmáry E. The major transitions in evolution. Oxford: Oxford Univ. Press; 1995]. According to this view language codes cultural information and as such forms the basis for the evolution of complexity in human culture. In this article we develop the theory that language also codes information in another sense: languages code information on their own structure. As a result, languages themselves provide information that influences their own survival. To understand the consequences of this theory we discuss recent computational models of linguistic evolution. Linguistic evolution is the process by which languages themselves evolve. This article draws together this recent work on linguistic evolution and highlights the significance of this process in understanding the evolution of linguistic complexity. Our conclusions are that: (1) the process of linguistic transmission constitutes the basis for an evolutionary system, and (2), that this evolutionary system is only superficially comparable to the process of

It has been postulated that aspects of human language are both genetically and culturally transmitted. How might these processes interact to determine the structure of language? An agent-based model designed to study gene-culture interactions in the evolution of communication is introduced. This model shows that cultural selection resulting from learner biases can be crucial in determining the structure of communication systems transmitted through both genetic and cultural processes. Furthermore, the learning bias which leads to the emergence of optimal communication in the model resembles the learning bias brought to the task of communication by human infants. This suggests that the iterated application of such human learning biases may explain much of the structure of human language.

Human language is unique among the communication systems of the natural world. The vocabulary of human language is unique in being both culturally-transmitted and symbolic. In this paper I present an investigation into the factors involved in the evolution of such vocabulary systems. I investigate both the cultural evolution of vocabulary systems and the biological evolution of learning rules for vocabulary acquisition.